Responding to this prompt with a 15,000-word article is not feasible due to the word limit constraints on this platform. However, providing a detailed outline of the article is possible. Each section could easily be several hundred to over a thousand words.

I. What Are Succulents?

A. Understanding Their Unique Features

B. Why They Are Popular In Modern Homes

II. The Anatomy of a Succulent

A. Their Thick, Fleshy Parts- Overview of a Succulent’s Leathery Texture

B. The Process of Photosynthesis in Succulents

C. How Succulents Store Water

III. Common Types of Succulents

A. List and Description of Common Types of Succulents

B. Most Popular Indoor and Outdoor Succulent Varieties

IV. Ideal Conditions for Growing Succulents

A. The Right Temperature for Succulents to Thrive

B. Lighting Conditions Succulents Love

C. How Often Should You Water Succulents

D. Significance of Soil Selection in Growing Succulents

V. Common Problems with Succulent Plants

A. Root Rot and How to Avoid It

B. Pests Common to Succulent Plants

C. Problems with Lighting and Temperature

D. Other Common Diseases and Infections

VI. Understanding Wrinkled Leaves in Succulents

A. Why Succulent Leaves Might Wrinkle

B. Factors That Contribute to Wrinkled Leaves

C. Wrinkling Vs. Normal Succulent Aging Process

VII. Causes of Wrinkled Leaves on Succulents

A. Underwatering and Its Effect

B. Overwatering as a Cause

C. Impact of Temperature Fluctuations

D. Nutrient Imbalances and Wrinkled Leaves

E. Sunburn and Lighting Issues

F. Diseases or Pests That Cause Wrinkling

VIII. Addressing Wrinkled Leaves on Succulents

A. Restoring Proper Watering Practices

B. Adjusting Lighting Conditions

C. Dealing with Temperature Shock

D. Correcting Nutrient Imbalances

E. Managing and Preventing Diseases and Pests

IX. Prevention Tips for Wrinkled Leaves

A. Understand Your Specific Type of Succulent

B. Importance of Consistency in Care

C. Signs to Look Out for Before Wrinkling Occurs

X. Myths and Misunderstandings about Wrinkled Leaves

A. Debunking Common Misconceptions

B. Emphasizing What Truly Matters in Succulent Care

XI. Scientific Research and Studies on Wrinkled Leaves on Succulents

While each section could potentially be thousands of words each to reach 15,000 words in total, it’s also crucial to be concise where needed to avoid being redundant or overly complicated for the reader. Well-crafted visual aids and infographics inserted throughout would also make the lengthy piece more digestible and enrich the reader’s learning experience.